Curriculum Vitae

PETER GREGORY MAY

  


   ADDRESS: Department of Biology
                    Stetson University
                    DeLand, FL 32723
                    (386) 822-8189
                    e-mail:
pmay@stetson.edu     
                   
   CURRENT POSITION: Professor



   EDUCATION :
 
      George Mason University, Fairfax, VA. B.S., Biology. May, 1977.

      George Mason University, Fairfax, VA. M.S., Biology. May, 1979. Thesis advisor: Dr. Carl
       Ernst. Thesis: "
Secondary succession and breeding bird community structure in the eastern   
       deciduous forest formation
".

      University of Florida, Gainesville, FL. Ph.D., Zoology.  May,1985. Advisors: Dr. Lincoln P.
       Brower and Dr. Carmine A. Lanciani. Dissertation: "
Foraging selectivity in adult butterflies:
       Morphological, physiological, and ecological factors affecting flower choice
".

TEACHING EXPERIENCE:

    University of Florida:
     Teaching assistant in Evolution, 1979, Functional Vertebrate Anatomy, 1980, General Biology I
      and II, 1980-84,Pollination Ecology, 1983.  Lecture in Cells and Organisms, 1980, Evolution,
      1986, Cells, Organisms, and Genetics, 1987-88. Coordinator of General Biology labs, 1984,
      coordinator and instructor for Laboratory in  Biological Sciences, 1985-86.

    Stetson University:     
      Lecture and laboratory instructor in: Man in the Biological  Community; Invertebrate
      Zoology; Comparative Vertebrate Anatomy; Man: From Cell to Organism; General
      Biology I; .General Biology II;   Entomology; Vertebrate  Zoology; Ornithology;  Special
      Topics in Ecological Research;  Junior seminar; Senior seminar; Experimental Biology II;
    
      Winter term courses:  Creationism vs. Evolution; The Role of Competition in  Ecology:   
      Communities and Ecosystems of Florida.
    
RESEARCH INTERESTS:

      General: Evolutionary and behavioral ecology of vertebrates and insects; plant reproductive
       strategies and evolutionary ecology. Food selection in animals. Herpetological natural history
       and ecology.

      Specific: Integration of larval and adult foraging strategies in holometabolous insects; resource
        vs. pollinator limitation of plant reproduction; mobbing behavior of passerine birds; population
        biology and natural history of reptiles and amphibians.

RESEARCH EXPERIENCE:

      Master's thesis: analysis of avian community structure.
    
      Dissertation: butterfly foraging energetics and behavior.

      Research assistant to Dr. Carmine Lanciani, 1980-81: effects of water mite parasites on
       notonectid bugs.
  
      Collaborator with Dr. Eugene Spears, Jr., 1982-84:  determinants and ecology of sexual
       expression in an andromonoecious plant, Passiflora incarnata.

       Stetson University, 1988- 1992:  Effects of sugar intake on fat storage in butterflies; Foodplant
        selectivity and its effects on  growth in lubber grasshoppers; Lipid storage patterns of   the zebra
        butterfly, Heliconius charitonius.
 
      Collaboration with Dr. Terence Farrell, 1992-1998:  Population and thermal ecology of pygmy
        rattlesnakes, Sistrurus miliarius barbouri, 1991-1995. Movement patterns and  demography of
        Florida box turtles, Terrapene carolina   bauri.  Crepuscular movement patterns of  aquatic
        snakes and frogs.

PHOTO CREDITS:

        
The Audubon Society Field Guide to Butterflies, Knopf; Snake of the United States and Canada,
        
Smithsonian; Snakes of the Southeast, University of Georgia; North American Box Turtles,
          University of Oklahoma;  Biodiversity in Lake Maracaibo, Shell Venezuela; Copperheads,
          Rosen; Cottonmouths, Rosen; Biology, LeBel; Science News; Rod and Reel Magazine;
          University of Georgia Research; Alabama Birdlife; CD Cover for Mockingbirds, Somerset
          CDs. Miscellaneous publications by Florida Freshwater Fish and Wildlife Conservation
          Commission; Pinellas County Environmental Management; Iowa Department of Natural
          Resources; Medical Action Communications;  Various museum exhibits, including Baylor
          University Natural History Museum,  Seneca Park Zoo Society, Nature Conservancy Blowing
          Rocks Preserve, National Park Service Point Lookout National Park.


 

PROFESSIONAL SERVICES :

      Referee for manuscripts and grant proposals from Auk, Oikos, NSF Postdoctoral Fellowship
       program, Functional Ecology, American Naturalist, Journal of Herpetology,  Chelonian
      Conservation and Biology, Journal of Wildlife Biology, Ecology, Biology of the Rattlesnakes.
   

HONORS, FELLOWSHIPS, GRANTS:

      Member, Phi Beta Kappa, Alpha Chi, Beta Beta Beta.
      Governor's fellowship, George Mason University, 1977.
      Graduate Council fellowship, University of Florida, 1979-1981.
      Research assistantship, Department of Zoology, University of Florida, 1983.
      Sigma Xi grant-in-aid of research: Energy maximization in foraging by Agraulis vanillae, Spring
       1982.
      O.C. Marsh fund grant, National Academy of Sciences: Energy maximization in foraging by
       Agraulis vanillae, Spring 1982.
      Nominee for University of Florida teaching assistant of the year, 1984.
      Stetson University summer research grants.  Summer, 1992, 1993, 1996, 1999.
      Volusia County Department of Environmental Management. Development of a management plan
       for Volusia County scrub habitats. $3500 (for student support), Summer 1992 (with T.M.
       Farrell).
      U.S.Department of Energy, Savannah River Ecology Laboratory. Equipment grant for snake
       research. $2100, Summer 1992. (with T.M. Farrell).
      Central Florida Herpetological Society. Population biology of pigmy rattlesnakes. December
        1993, $300; May 1994, $1000; June 1996, $2200.; $1000, June 1997 (with T.M. Farrell).
      Explorers Club of Central Florida. Population biology of pigmy rattlesnakes. May 1994, $1500.
       (with T.M. Farrell).
      AVID, Corp., Equipment grant for pigmy rattlesnake study. August, 1994, $2100. (with T.M.
        Farrell).
      Suncoast Herpetological Society, May 1996, $150. 1997, $100. (with T.M. Farrell).
      Gainesville Herpetological Society, May 1996, $250. 1997, $300. (with T.M. Farrell).
      Everglades Herpetological Society, 1997, $200. (with T.M. Farrell).
      Volusia County Herpetological Society, 1997, $150. (with T.M. Farrell).
      Homer and Dolly Hand Faculty Research Award, May 2000. 

PAPERS PRESENTED AT MEETINGS:

    Florida Academy of Sciences, St, Petersburg, FL, March 1993:
        T.M. Farrell, P.G. May and M.A. Pilgrim.  Reproduction in the dusky pigmy rattlesnake,
         Sistrurus miliarius  barbouri).
        J.N. Sourial, T.M. Farrell and P.G. May. The impact of PIT tagging on the growth and
         movement of the dusky pigmy rattlesnake (Sistrurus miliarius barbouri).
       Pilgrim, M.A., P.G. May and T.M. Farrell.  Growth in juvenile dusky pigmy rattlesnakes,
        Sistrurus miliarius barbouri.
       Everett, J.W., S.C. Jemison, P.G. May, and T.M.  Farrell.  The seasonal activity and thermal
         biology of the dusky pigmy rattlesnake, Sistrurus miliarius  barbouri.
      May, P.G. Effects of sugar type on meal size and lipid  storage in the gulf fritillary,
Agraulis
        vanillae
(Lepidoptera: Nymphalidae).

      American Society of Ichthyologists and Herpetologists, The Herpetologists League,
Austin,
      TX
, May-June 1993:
       May, P.G. and T.M. Farrell. The seasonal activity and  thermal biology of the dusky pigmy
        rattlesnake,
Sistrurus miliarius barbouri.
      
Farrell, T.M., P.G. May and M.A. Pilgrim.  Reproduction in  female dusky pigmy rattlesnakes,
        Sistrurus miliarius barbouri.
 
     Florida Academy of Sciences, Tallahassee, FL., March 1994:
        Pilgrim, M.A., T.M. Farrell and P.G. May.  Annual variation in reproductive parameters in a
        central Florida population of rattlesnakes, Sistrurus miliarius.
       Bishop, L.A. , T.M. Farrell, and P.G. May. Sexual dimorphism in the rattlesnake
Sistrurus
        miliarius
barbouri.

     Herpetologists' League, Society for the Study of Amphibians and Reptiles, Athens, GA.,
       July- August 1994:
       May, P.G. and T.M. Farrell. Variability in winter activity of dusky pigmy rattlesnakes (
Sistrurus
        miliarius
 barbouri).
       Farrell, T.M. and P.G. May.  Are potential fecundity costs a major component of reproductive
        effort in the rattlesnake Sistrurus miliarius?
       Bishop, L.A. , Farrell, T.M. and P.G. May.  Sexual dimorphism in the rattlesnake
Sistrurus
        miliarius
.

     All-Florida Herpetological Conference, April 22, 1995:
       May, P.G. and T.M. Farrell. Population ecology of pygmy  rattlesnakes in central Florida.
        Invited presentation.

     Society for the Study of Amphibians and Reptiles, BooneN.C., August 1995:
       May, P.G. and T.M. Farrell. Seasonal patterns of abundance and activity in dusky pigmy
        rattlesnakes (Sistrurus miliarius barbouri).
       Farrell, T.M., P.G. May and M.A. Pilgrim.  What causes offspring size variation among
        rattlesnake litters?

     Society for the Study of Amphibians and Reptiles, LawrenceKansas, July 1996:
       May, P.G. and T.M. Farrell. Recapture frequency in pigmy  rattlesnakes and its association with
        body condition and movement.
       Farrell, T.M. and P.G. May. Why did rattlesnakes evolve  rattles? Evidence from recent studies
        of  pigmy rattlesnakes (Sistrurus miliarius).
       Pilgrim, M.A., Farrell, T.M. and P.G. May.  Annual variation in the reproductive traits of the
        rattlesnake  Sistrurus miliarius.

    La Selva Biological Station, Organization for Tropical  Studies, Puerto Viejo de Sarapiqui,
       Costa  Rica. December, 1996:
         May, P.G. and T.M. Farrell.  Population  ecology of pigmy rattlesnakes in central Florida.
          Invited presentation.

   
American Society of Ichthyologists and Herpetologists, Herpetologists’ League, Society for
      the  Study of Reptiles and Amphibians
, Seattle, Washington, June-July, 1997:
        Farrell, T.M., M.A. Pilgrim and P.G. May.  Population structure, activity and sexual
         dimorphism in a central Florida population of box turtle, Terrapene carolina.
       Heulett, S.T., T.M. Farrell and P.G. May.  Caudal luring and tail coloration in snakes: An
         experimental and comparative approach to the study of evolutionary relationships.

 

      Florida Academy of Sciences, Orlando, FL. April, 1998.

        Cheatwood, J.L.,  P.G. May, and T.M. Farrell.  Coccidian parasites in a central Florida
          population of pigmy rattlesnakes, Sistrurus miliarius barbouri
       Hendrie, A.M.,  T.M. Farrell, and P.G. May.  The effects of temperature and season on the
         activity of the Florida box turtle.
       LaBram, J.A., T.M. Farrell and P.G. May. The home range areaof the Florida Box turtle,
         Terrapene carolina, in central Florida.
       Siefkas, J., T.M. Farrell, and P.G. May.  Infection by Mycoplasma agassizii in a box turtle and
         its implications for conservation biology.

 

     American Society of Ichthyologists and Herpetologists, Herpetologists’ League, Society for  
     the Study of  Reptiles and Amphibians,
Guelph, Ontario, July 1998.
       Farrell, T.M. and P.G. May.  The impact of fecundity on parturition date in the rattlesnake,
        Sistrurus miliarius.
       May, P.G. and T.M. Farrell. Maternal attendance and post-parturient behaviors of neonate pigmy
        rattlesnakes.
       Pilgrim, M.A., T.M.  Farrell and P.G. May. Variation in offspring characteristics of
Sistrurus
        miliarius, Sistrurus  catenatus, Thamnophis sirtalis, Thamnophis proximus,
and
Nerodia
        rhombifer.

         
PUBLICATIONS:

       Lanciani, C.A. and P.G. May. 1982. Parasite mediated reductions in the growth of nymphal
         backswimmers. Parasitology 85: 1-7.

      May, P.G. 1982. Secondary succession and breeding bird community structure: patterns of
         resource utilization.  Oecologia 55: 208-216.

      May, P.G. 1984. Avian reproductive output in early and late  successional habitats. Oikos 43:
       277- 81.

      May, P.G. 1985. Nectar uptake rates and optimal nectar concentrations in two butterfly species.
       Oecologia 66: 381-386.

      May, P.G. 1986. A simple method for measuring nectar uptake rates. J. Lepidopterists' Soc. 39:
       53-55.

      May, P.G. and E.E. Spears, Jr. 1988. Andromonoecy and the regulation of sexual expression in
       Passiflora  incarnata. American J. Botany 75(12): 1830-1841.

      Spears, E.E., Jr. and P.G. May. 1988. The effect of  herbivory on sexual expression and fruitset
       in  Passiflora  incarnata. American J. Botany 75(12): 1830-1841.

      May, P.G. 1988. Determinants of foraging profitability in two species of nectarivorous
        butterflies. Ecological Entomology, 13: 171-184.

      May, P.G. 1992.  Flower selection and the dynamics of lipid reserves in two nectarivorous
        butterflies. Ecology 73: 2181-2191.

      May, P.G. 1993. Effect of sugar type on food intake and lipid dynamics in adult
Agraulis
        vanillae
(Lepidoptera, Nymphalidae). Journal of the Lepidopterists' Society  47:279-290.

      May, P.G., and T.M. Farrell. 1994. Researching the real serpent: Studies of
        pigmy rattlesnakes. Newsletter of the League of Florida Herpetological Societies, May 1994.

      Farrell, T.M., P.G. May and M.A. Pilgrim. 1995. Reproduction in the rattlesnake,
Sistrurus
       miliarius barbouri
, in Central Florida.  Journal of Herpetology 29:21-27.

      Jemison, S.C., Bishop, L.A., P.G. May and T.M. Farrell. 1995. The impact of PIT-tags on
       growth  and movement of the rattlesnake, Sistrurus miliarius. Journal of Herpetology 29: 129-
       132.

      Bishop, L.A. , T.M. Farrell and P.G. May. 1996. Sexual dimorphism in the rattlesnake
Sistrurus
       barbouri
.  Herpetologica 52: 360-364.

      Aycrigg, A.D., T.M. Farrell, and P.G. May.  1996.  Natural History Note: Seminatrix pygaea:
        Predation.  Herpetological Review 27: 84.

      May, P.G., Farrell, T.M., Pilgrim, M.A., Heulett, S.T., Bishop, L.A., Spence, D.J., Rabatsky,
       A.R., M.C. Campbell, Aycrigg, A.D., and W.E. Richardson II.  1996. The seasonal abundance
       and activity of a rattlesnake (Sistrurus  miliarius barbouri) in central Florida. Copeia 1996: 389-
       401.

      May, P.G., S.T. Heulett, T.M. Farrell, and M.A. Pilgrim.1997. Live Fast, Love Hard, and Die
       Young: The Ecology of  Pigmy Rattlesnakes.  Reptile & Amphibian magazine 45: 36-49. 

     Pilgrim, M.A., T.M. Farrell and P.G. May. 1997.  Population structure, activity, and sexual
       dimorphism  in a central  Florida population of Terrapene carolina. Chelonian Conservation and
       Biology, 2: 483-488.

     May, P.G. and T.M. Farrell. 1997.  Death from above: Adventures with arboreal snakes.
        Reptiles, 5: 76-83.

     Aycrigg, A.D., T.M. Farrell, and P.G. May. 1997. SOS: Sounds of Survival.  Reptile &
       Amphibian,  52: 56- 63.

     May, P.G. and T.M. Farrell. 1998.  Florida's flatwoods cottonmouths.  Reptile & Amphibian
       magazine, 56:18-24.

     Roth, E.R., P.G. May, and T.M. Farrell. 1999. Pigmy rattlesnakes use prey-derived odor cues to
      select  foraging sites.  Copeia 1999: 772-774.

     Rowe M.P., T.M. Farrell, and P.G. May. 2002.  Rattle Loss in Pigmy Rattlesnakes (
Sistrurus
      
 miliarius): Causes, Consequences, and Implications for Rattle Function and Evolution. pp. 385-
        404 In BIOLOGY OF THE VIPERS, edited by: Gordon W. Schuett, Mats Höggren, Michael E.
       Douglas, and Harry W. Greene. Eagle Mountain Publishing, LC. Eagle Mountain, Utah.

 

    Greene, H.W., P.G. May, D.L. Hardy, Sr., J.M. Sciturro, and T.M. Farrell. 2002.  Parental
       Behavior by Vipers.  pp. 179-206 In BIOLOGY OF THE VIPERS, edited by: Gordon W.
       Schuett, Mats Höggren, Michael  E. Douglas, and Harry W. Greene Eagle Mountain Publishing,
       LC. Eagle Mountain, Utah.


    Cheatwood,  J.L., E. R. Jacobson, P.G. May,  T.E. Farrell B. L. Homer,  D. A. Samuelson, and  J.
       W.  Kimbrough. 2003.  An outbreak of fungal dermatitis and stomatitis in a free-ranging
        population of pigmy rattlesnakes (Sistrurus miliarius  barbouri) in Florida  Journal of Wildlife
        Disease 39: 329-337.

 

      Glaudas, X., T.M. Farrell, and P.G. May. 2005 .  Defensive Behavior of Free-Ranging Pygmy
         Rattlesnakes (Sistrurus miliarius).   Copeia.
2005: 196–200 .

 

   Farrell, T.M., D.H.. Dodd, Jr., and P.G. May.  2006.  Terrapene carolina – eastern box turtle.  pp.

         235-248 in Biology and Conservation of Florida Turtles, P.A. Meylan (ed.).  Chelonian
         Research Monographs No. 3, 376.

 

     Farrell, T. M., M.A. Pilgrim and P.G. May.  2008.  Annual variation in neonate recruitment in a
          Florida population of the Pigmy Rattlesnake, Sistrurus miliarius. pp.
257-264
in W.K. Hayes,
          K.R. Beaman, M.D. Cardwell, and S.P. Bush (eds.), The Biology of Rattlesnakes.  Loma Linda
          University Press, Loma Linda, California.

   

   Farrell, T.M, P.G. May, and M.A. Pilgrim.  2009. Repeatability of Female Reproductive Traits in a  
          Population of Pigmy Rattlesnakes (Sistrurus miliarius).  Journal of Herpetology  43: 332–335.

 

    Farrell, T.M., M.A. Pilgrim, W.B. Blihovde, and P.G. May.  2011. The Herpetofauna of Lake
          Woodruff National Wildlife Refuge, Florida.  Southeastern Naturalist 10(4):647-658.

 

   Pilgrim, M.A., T. M. Farrell, Peter G. May, M.R. Vollman, and R.A. Seigel. 2011. Secondary Sex
            Ratios in Six Snake Species.  Copeia 2011, No. 4, 553–558.

 

   Farrell, T.M., P.G. May, and P.T. Andreadis.  2011. Experimental manipulation of tail color does not
            affect foraging success in a caudal luring rattlesnake. Journal of Herpetology 45(3):291-293.

 

   May, P.G. and T.M. Farrell. In press.  Growth Patterns of Dusky Pigmy Rattlesnakes (Sistrurus miliarius
            barbouri
) from Central Florida. Herpetological Monographs.